Williams, who lost in the final to sister Serena last year, was stunned by Swiss star Belinda Bencic, fresh from winning the Hopman Cup with Roger Federer.
The seven-time Grand Slam winner, in her 77th major, struggled against a player who had never before beaten her to go down 6-3, 7-5 and deprive the tournament of one of its biggest names.
It is the first time since 1997 that there will be neither of the Williams sisters in the second round, with Serena not playing after giving birth to her first child.
“I would like to have played someone easier,” said Bencic.
“When I was a little girl watching on TV I would never imagine I would play them (the Williams).”
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